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The claim in the American Rifleman magazine was that Gussler says the octagonal shape is an exterior representation that the bore is rifled. That struck me as quite possibly being an unverified myth for the sake of impressing the tourists. Personally, I am far less expert than many on this forum. But, I have been at this game for a half century and have visited  more than just a few gun museums. Viewing the long guns in those museums would have illustrated this notion. I have never seen, or heard, it from any other source.